Tentacruel in depth
Eighty tentacles and a Portuguese man o' war
The Japanese is descriptive and little more: ドククラゲ combines 毒 (poison) with 水母 (jellyfish). The English name put a bit more work into it, combining tentacle with cruel, and during development another name was considered, Man-O-War, pointing directly to the Portuguese man o' war. The reference fits better than a generic jellyfish: the Portuguese man o' war isn't an animal but a colony of organisms, with stinging tentacles that can measure dozens of metres and a floating bladder at the surface. Hence the translucent body, the independent tentacles and the habit of appearing in schools.
The red orbs have no biological equivalent. They're a design addition that serves as a signal — they shine when the creature gets agitated — and as an excuse for the ultrasonic blast described in the Pokédex entries.
What the Special Attack split did to it
In the first generation, it had a single Special stat of 120, and that number governed both the power of its special attacks and its resistance to special opponents. With Water and Poison on top, it was a Pokémon that hit hard and took special hits hard at the same time.
The second generation split that stat in two, and the distribution was unkind: 120 stayed in Special Defense, while Special Attack fell to 80. Suddenly, it stopped being an offensive threat and became what it still is today, a fast bulky Pokémon with mediocre damage output. Abilities arrived later, in the third generation, and Rain Dish later still as a hidden ability.
A resemblance that isn't a relationship
Paldea introduced Toedscruel, with the same bell-shaped silhouette and tentacles, the same general feel and no evolutionary connection: it is a convergent species of Ground and Grass type, a fungus that imitates a jellyfish. The joke works because the original was never exactly a recognisable marine animal.
In the water, the useful comparison is different. Toxapex fills the same role as a poisonous nuisance with much better defences and ridiculous Speed; Starmie shares Rapid Spin and high Speed but can actually hit hard. Its own niche is the 120 Special Defense combined with its ability to absorb Toxic Spikes, which neither of them offers.
How to play it in a normal playthrough
The level-up movepool is stingy and comes late. Bubble Beam at 24 and Water Pulse at 16 have to carry the entire middle of the game, because Surf doesn't appear until 46, Sludge Wave until 52 and Hydro Pump until 58. In between are two status moves that make up for its weak point: Acid Armor at 34 raises Defense by two stages, precisely the stat sitting at 65, while Screech at 20 drops the opponent's by another two.
Supersonic, at 12, is the standard confusion move with unreliable accuracy. And Reflect Type, available from level 1, copies the opponent's type, something that in a game against the AI is more useful for curiosity than for winning. The practical route goes through the TMs: Sludge Bomb, Ice Beam and Toxic cover almost everything a Water-type needs to cover.