Illumise in depth
Luciferase and Charleston
The English and international name crosses illuminate with luciferase, the enzyme that produces bioluminescence in real fireflies; the Japanese イルミーゼ comes from the same place. The biological basis is a lampyrid, just as with Volbeat, but the design does not stop at the insect. The purple bob, the black body that looks like a jacket and the rounded white collar point to 1920s fashion and the silhouette of the flappers. It is a party outfit put on a firefly, which fits a creature whose job is to organise night-time spectacles.
Thirty points that arrived late
For four generations, from the third through the sixth, its base stat total was 400. In the seventh, it received a ten-point increase in Defense and ten in Special Defense, giving it the 75 and 85 shown today. Volbeat received exactly the same adjustment. It was a small change and arrived late: it barely changes what Illumise can do, because the problem was never taking a hit, but giving one back.
Its availability has also been narrow. On debut, it was limited to Hoenn's Route 117, and in the fourth generation it reappeared on Sinnoh's Route 229. In Pokémon GO it is regional: it appears in the Americas, Africa and the Middle East, while Volbeat occupies the rest of the map, which is the most literal way possible to split up a pair.
How to use it in a normal playthrough
Outside a purpose-built team, Illumise's value lies in the order in which it gets its moves. Sweet Scent at 5 and Charm at 9 make it a comfortable companion from very early on; Moonlight at 19 gives it self-recovery, and Wish at 22 heals whoever takes its slot two turns later. Flatter at 29 confuses the opponent in exchange for raising their Special Attack, and against a physical attacker that is a reasonable trade. Encore at 26 is the move that gets the most mileage out of a slow Pokémon against an opponent who has just used something useless.
What it does not have is power. Its TM list is long and tempting—Thunder, Solar Beam, Shadow Ball, Dazzling Gleam, Focus Punch—and almost all of it runs through stats that do not measure up. Compared with Butterfree, which fills a similar Bug-type support niche, Illumise loses in Special Attack but wins in Special Defense and in Prankster priority, which Butterfree does not have.
A pair that never quite shared a screen
Volbeat and Illumise are among the few cases in which the game splits one species into two Pokédex numbers by sex instead of into two forms of the same Pokémon, with the result that each gets its own entry, its own data line and its own capture slot. Illumise's Pokédex entries talk about Volbeat; Volbeat's talk about Illumise. Neither explains itself on its own, and both are mediocre separately.